B.  Gantenbein, Fet, V., Largiader, C., e Scholl, A., «First DNA phylogeny of Euscorpius Thorell, 1876 (Scorpiones: Euscorpiidae) and its bearing on taxonomy and biogeography of this genus», Biogeographica (Paris), vol 75(2), pagg 49-65, 1999.